Index

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Page numbers in bold refer to illustrations; page numbers in italics refer to information contained within tables.

Aberdeen, 58, 77-8

Aberdeenshire, 147

Abernethy, 119-22, 164, 288, 329, 406-7, 410

Aboyne, 138

afforestation, 113, 118-22, 391, 400, 405-6

Alaska, 1, 31-3, 39, 43, 68-9, 76-82, 86-8, 174, 184, 197-9, 204, 206, 210, 215, 231, 250-1, 257, 300, 304-5, 365, 381

Eagle Creek, 91-3, 95, 98, 194, 196, 209

Porcupine Creek, 37, 225, 233

alder, Alnus glutinosa, 113, 267, 270

Aleutian Islands, 32-3, 68, 76-7, 80, 82-3, 99, 222, 240-3, 320

Alps, 33, 67, 82, 109, 412

Bavarian, 108, 163-4, 167

French, 385-6

Italian, 91

Swiss, 87-9, 91, 95, 224

altitude, 39, 80-1, 90-1, 95

America, 31, 77, 79, 87-8, 134, 174, 260, 381

North, 1, 4, 8-11, 56, 65, 78, 303, 373, 384, 407

Angus, 398

Argyll, 110, 131, 148

Argyllshire, 147

Arran, 77, 277

Asia, 146, 303, 305

aspen, Populus tremula, 157, 148, 270, 308, 409

Atholl estate, 124, 125

Austria, 147

Avalonia, 259

avens, mountain, Dryas octopetala, 84

azalea, mountain, Loiseleuria procumbens, 81

Baffin Island, 33, 81, 87, 243, 266-7

Balkans, 146

Ballochbuie, 151, 407

Balmoral, 147

Baltica, 259

Baltic States, 32, 49

Banchory, 409

Bear Island, 77

beck, 20, 176, 212, 221-2, 384

bedrock, 260, 284-5, 288

effect on soil fertility, 46-8, 91, 261, 262, 272, 350

bedstraw, heath, Galium saxatile, 309

beech, Fagus sylvatica, 113, 155

Belgium, 33

Ben Lomond, 77

Beringia, 1, 2, 4, 8, 146

Bering Island, 76

bilberry, see blaeberry

bill, 253

Biodiversity Action Plan, 131

birch, Betula, 265-7, 276-7, 398, 399, 406, 409

habitat, 38, 89, 148, 270

in diet, 11, 83-4, 94, 113-4, 287, 291, 303-5, 308

bistort, Alpine, Polygonum viviparum, 84, 88, 94, 99, 183

blackcock, see grouse, black

blackgame, see grouse, black

blaeberry, Vaccinium myrtillus, 95, 110-13, 116, 119, 149, 151-4, 172, 304, 413-14

habitat, 81, 164, 270, 281, 288, 377-8, 410

in diet, 10, 38-9, 83-4, 86, 113, 115, 148, 155-6, 291, 300, 301-2, 305, 312

influence of bedrock and soil on, 261, 277, 284-6, 407

Blelack, 410

blueberry, Vaccinium (in North America), 39-40

Bohemia, 333

bracken, Pteridium aquilinum, 156, 396, 397, 407, 414

Braemar, 350

Brecon, 109

Bristol Channel, 269

Buccleuch estate, 124, 125

Buchan, 147

Bulgaria, 76

bursa of Fabricius, 237

buzzard, Buteo buteo, 66, 322, 324, 325, 332

Bylot Island, 87

Cabrach, 77

Cairn Gorm, 98, 100, 119, 201, 218, 318, 368, 406, 412

Cairngorms, 33, 44, 52, 54, 78, 81, 87, 89, 91, 95, 190, 203, 211, 240-1, 277, 311, 318, 320, 345, 350, 353-4

National Park, 154, 402

Cairnwell, 50, 52, 83-4, 87, 90, 91-5, 98-9, 205, 216, 221, 228, 240, 285, 311, 345, 350-5, 372, 384, 411

Caithness, 77, 267, 270, 281

Caledonian mountains, 260

Caledonian pinewood, 148, 149, 151, 168, 259, 377, 406-7, 408, 414

camouflage, 28, 74, 96, 100, 196, 238-40, 317

Canada, 32-3, 35, 50, 54, 62, 67, 78, 88-9, 91, 93, 95, 98, 100, 184, 192, 194-5, 234, 238, 247, 260, 277, 320, 337 Baffin Island, 33, 81, 87, 243, 266-7

Bathurst Island, 222

British Columbia, 39, 76-7, 82, 257

Chilkat Pass, 43, 49, 53, 57-8, 202, 208, 213, 215, 224, 227-8, 231, 233

Bylot Island, 87

Churchill, 196

Ellesmere Island, 86

Hudson Bay, 269

Labrador, 76-7, 87, 413

Newfoundland, 31, 39, 57, 77, 187, 209, 224, 239, 365-6, 384

Brunette Island, 49-50, 53

Northwest Territories, 88, 93, 242, 413

Ungava Peninsula, 81, 87, 94, 213, 216, 242, 413

Cantabrian mountains, 146, 148

capercaillie, Tetrao urogallus, 6, 16, 134-7, 290

behaviour, 137-8, 139-43, 144-5, 314

calls, 138, 175

feeding, 182-4, 303

parental care, 191-3

preening, 184, 187

resting, 187

roosting, 188-9

in snow, 198-9, 203-4, 207-8

social, 189-90

territorial, 173-4

diet, 44, 155-7, 291, 307-10, 313, 315

distribution, 147-8, 157

enemies, 152, 153, 168, 169-71, 318-46, 400, 409, 413-14

growth, 157

habitat, 120, 122, 148, 149-51, 152, 154, 172, 287-8, 410

fragmentation, 165-7, 376-7, 379

improvement, 407

metabolism, 157

moult, 158

movements, 162-5

plumage, 136, 235-7

population:

decline, 134, 166-8, 169-1, 331-2

density, 120, 122, 133-4, 145-6, 147, 166-8, 319

fluctuation, 52-4, 165, 372

size, 147-8

reproduction:

breeding success, 163, 168, 172, 313-5, 335

chicks, 144, 152, 154-7, 161, 163, 168, 180, 181, 191-6

eggs, 26, 159, 161

lek, 108, 137-8, 139-46, 151-2

nests, 153, 159-60

shooting, 133-4, 333, 385-6

size, 135-7, 157, 314

survival, 157-8, 161, 314

weight, 135

capercaillie, black-billed, Tetrao parvirostris, 2, 8, 10

caterpillar, 129, 156, 161, 164, 313, 315, 319, 396

Catterick, 363

cedar, Cedrus, 202

Channel Islands, 269

Chapman, Abel, 384

chickweed, Stellaria, 38, 84

China, 384

claws, 253-4

climate, 9, 12, 21, 91, 148, 167, 277-9, 289, 354

change, 11, 55, 66, 100-1, 132, 154, 202, 283-4, 345-6, 413-14

effect on egg development, 58, 95

effect on habitat, 34, 46, 79, 80, 263-5, 266-7, 268-70, 276, 288

effect on parasites, 336, 345-6

effect on plumage , 238-9, 247, 258

Coilacriech, 164

combs, 255-8

compensation, 385-7

Corndavon, 53

Cornwall, 109

cotton-grass, Eriophorum, 281, 307

effect on breeding success, 60, 64, 94, 99, 155-6, 312

hare’s-tail, E. vaginatum, 38-9

in diet, 34, 48, 57, 84, 114, 164, 183, 287, 308-9

courtship, 179-80, 232

Cowal Peninsula, 147

cowberry, Vaccinium vitis-idaea, 82-3

cranberry, Vaccinium oxycoccus, 407

crow (carrion, Corvus corone; hooded, C cornix), 65, 100, 119, 324, 325, 329, 368-9, 412

(Note: Taxonomists have long considered the carrion and hooded crow as varieties of one species, but recently decided that there were two. Because the two interbreed in much of Asia and Europe including Scotland, and produce many hybrids of varying colour from black birds typical of carrion crows to grey ones typical of hooded or grey crows, we refer in the text simply to crows.)

crowberry, Empetrum, 82-6, 85, 115, 265-7, 281, 285-6, 291, 304-5, 312, 414

Cumbria, 283

Czechoslovakia, 412-13

Dartmoor, 33, 48

Deeside, 52, 127, 138, 167, 187, 240, 280, 287, 332, 407-10

deforestation, 34, 268-9, 277-9, 281-2, 284, 288, 344, 376

Denmark, 147

Derbyshire, 109

Derry Cairngorm, 78, 91-100, 223, 233, 240, 310

Deveronside, 409

Devon, 109, 265

diet, 154, 158-9, 300, 302, 304-5, 308-9, 312, 314, 316

digestion, 292-303, 293, 295-6, 301-2, 316

disease:

grouse, see threadworm, caecal louping ill, 342-6, 374-5, 379, 398, 400

dispersal, 4, 44-5, 88, 123, 152, 162-7, 227, 367

distraction display, 144, 191-2, 194-6

distribution, 8, 11-12

disturbance, 100, 108, 152, 193-4, 221, 315, 410

Donegal, 77

Donside, 167

Dorset, 109

dropping, 38, 187-8, 199, 221, 292, 294, 296

caecal, 292, 294, 296

clocker, 20, 60, 294, 297

Dumfries and Galloway, 131

Dunkeld, 147

Durham, 336

eagle, golden, Aquila chrysaëtos, 99-100, 191, 322, 324, 326, 330, 378,

white-tailed, Haliaeëtus albicilla, 322

Easter Ross, 167

economics, 403, 404-5

Edinburgh, 265

Edinglassie, 363

Ellesmere Island, 86

elm, Ulmus, 266-7

Eurasia, 1, 4, 31, 36, 49-50, 61, 65, 109, 187

Europe, 133, 147, 152, 166, 260, 269, 277, 317, 333, 336, 348, 385, 412-13

evolution, 1, 2-3, 4, 8, 9, 12, 315, 344

Exmoor, 33, 48

extinction, 11, 147, 172

Faeroes, 33, 77

falcon, peregrine, Falco peregrinus, 99, 322, 325, 328, 402

feathers, 251, 252-3, 258

felling, 133, 147, 154, 168

fences, 117-18, 152-3, 158, 168, 169, 171, 407, 409

Fennoscandia, 32, 166, 305, 319, 333, 368, 373, 376, 385

Finland, 33, 49, 52-4, 76, 111, 147, 165, 190, 198, 204-9, 270, 333-5, 372-3, 376

fir, 10

Douglas, Pseudotsuga menziesii, 155

Firth of Clyde, 265

flight, 28, 187

forest grouse, 2-3, 4, 8, 10, 137, 372

forestry, 113, 148, 154, 409

Forestry Commission, 133, 409-10

Forvie, 53, 366

fox, Arctic, Alopex lagopus, 99, 320

red, Vulpes vulpes, 41, 42, 59, 65, 99-100, 158, 170, 178, 192, 202, 209, 282, 318, 319, 320, 321, 324, 325, 329, 332-4, 378

France, 279, 381

Franz Josef Land, 77

Freuchen, Peter, 381

Galloway, 267-8, 282

Gannochy, 398

geological processes, 259, 260-1, 262, 270-7, 271-3, 274-5

Germany, 33

glaciation, 9, 265-6, 276

Glas-choille, 57, 218-20, 227, 233

Glen Affric, 269

Glenamoy, 29, 46, 60, 64, 175, 218, 222, 232-3, 278

Glen Clunie, 395

Glen Derry, 262

Glen Dye, 189, 363

Glen Esk, 39, 43, 56-62, 195, 217-19, 226-33, 229, 279, 338-40, 374, 383, 390, 395-6

Glen Feshie, 278

Glen Gairn, 399

Glen Livet, 77

Glen More, 406, 410

Glen Tanar, 126-30, 157, 162-4, 171, 230, 233, 359, 407, 410

glutton (wolverine), Gulo gulo, 320, 321

Golspie, 147

Goshawk, Accipiter gentilis, 318, 322, 324, 329

Grampian, 110, 262

grazing, 388-9, 405

effects on grouse populations, 66, 119, 120

effects on food availability, 60, 64, 78, 84, 94, 99, 152, 231, 307, 316

effects on habitat, 50, 100, 112, 115, 121, 126, 132-3, 151, 154, 277-8, 282-4, 286-8, 308, 330, 376-7, 393, 394, 395-6, 404, 407, 408, 414

management, 116-17

Greenland, 25, 33, 43, 67, 76-8, 81-9, 183, 187, 196, 203-4, 208, 210, 243, 260

greyhen, see grouse, black

grit, 183-4, 292, 295

grouse:

black, Lyrurus tetrix, 7, 114, 165, 254-5, 310

behaviour, 314

calls, 104, 175

feeding, 182-4, 303

parental care, 191-3

preening, 184, 187

resting, 187

roosting, 188-9, 206

in snow, 198-9, 202, 204, 207-S, 209

sexual, 141

social, 189-90

territorial, 173-4, 178, 211

diet, 44, 113-15, 129, 155, 291, 307-9

distribution, 109-10

enemies, 126, 131, 318-46, 400, 413-14

growth, 314

habitat, 101, 110-13, 111-12, 115-18, 120, 123, 132, 259, 270, 286-8, 376-9, 387, 409-10

improvement, 115-8, 121, 122

movements, 122-4, 162

plumage, 101-3, 235-7, 248-9, 258

population:

decline, 124-27, 125

density, 52-4, 119-24, 120, 123, 126, 348-9, 381

fluctuation, 124, 125, 372

size, 110

reproduction:

breeding success, 119, 122, 123, 127, 128-9, 130, 314

chicks, 102, 114, 128, 129-31, 157, 287

eggs, 26, 129

lek, 103-7, 108-9, 120, 123, 196

nests, 128-9, 255, 287

shooting, 385-6

size, 314

survival, 129-31, 130, 157, 314

weight, 101-2

blue, Dendragapus obscurus, 3, 8, 11, 162, 174, 303

Caucasian black, Lyrurus mlokosiewiczi, 2, 8, 9, 11-12, 109

Chinese, Bonasa sewerzowi, 2, 4, 8, 9, 12

dusky, Dendragapus obscurus, 3, 8

Gunnison sage, Centrocercus minimus, 3, 8, 12

hazel, Bonasa bonasia, 52-3, 165, 187, 207-8, 270, 321-4, 372, 385

red, Lagopus lagopus scoticus, 5, 30, 39, 44, 177, 289, 291, 328, 390

behaviour, 21, 28, 31, 41, 314

calls, 174-9, 175, 211-12, 218-22, 226

feeding, 180-3, 184-5

parental care, 191-6

predator avoidance, 190

preening, 184, 186

resting, 187

roosting, 181, 188-9

in snow, 197-210, 198, 205, 207-8

sexual, 179-80

social, 178, 180, 189-90

territorial, 173-8, 211-16, 214, 220-7, 232-4, 244, 342

effects on population density, 358-9, 360, 363, 364-5

kinship, 361, 362, 364-6, 371, 373, 375

diet, 38-40, 44, 291, 300-3, 301-2, 305, 307-10, 313

distribution, 32, 33

enemies, 65, 66, 190-1, 318-46, 368, 374, 378, 400, 412-14

growth, 27, 29, 314

habitat, 34, 35-7, 36, 46-8, 47, 52, 280-4, 377, 379, 394-6, 403, 406

metabolism, 40-2, 41, 292

movements, 43-6, 367

plumage, 21, 23, 31, 59, 235-7, 243—7, 248-9, 250, 256

population:

decline, 233, 339-41, 346

density, 36, 46-8, 51, 52-4, 55, 212-13, 222-4, 226, 342, 346, 348, 349, 381

fluctuations, 339-41, 346, 350-3, 351-2, 355, 366, 367-74, 370, 379, 385-7

size, 33

reproduction:

breeding success, 62-4, 63, 195, 230-2, 336, 348, 366

chicks, 27-8, 42, 226

eggs, 26, 27, 58-60, 61, 309

incubation, 41, 42, 58-9, 194-5, 226, 244, 250

nests, 58, 59

sex ratio in spring, 57

shooting, 55, 56, 62, 193, 329-30, 368, 381-7, 382-3, 397-8, 403, 414

size, 23, 314

survival, 56, 57, 340, 366, 387

territory, 211-12, 216-17, 218, 220, 223-4, 226-8, 230-2, 234, 367

weight, 23

ruffed, Bonasa umbellus, 2, 4, 8, 52, 137, 162, 174, 202, 207, 308, 337, 385

sage, Centrocercus urophasianus, 3, 8

sharp-tailed, Tympanuchus phasianellus, 3, 8

Siberian, Falcipennis falcipennis, 2, 8, 10, 12

sooty, Dendragapus fuliginosus, 3, 8

spruce, Falcipennis canadensis, 2, 4, 8, 10, 162, 184

white, see ptarmigan, Lagopus

willow, see willow ptarmigan, Lagopus lagopus

wood, see capercaillie, Tetrao

growth, 29, 42, 236, 312, 315

gyrfalcon, Falco rusticolus, 93, 94, 242, 319, 322, 373

habitat, 9-11, 276, 414

effect of climate on, 263-5, 266-8, 270

effect on grouse, 261

fragmentation, 10, 50, 66, 147, 319, 333, 347, 368-9

improvement, 168, 170

management, 55, 387-400, 388-9, 390-2, 392-3, 394-7, 399, 403-10, 414

man’s effect on, 268-9, 277, 278-80, 387, 388-9, 390-2, 398, 400, 409-14

Hampshire, 109

handicap principle, 241-2

hare, mountain, Lepus timidus, 66, 318, 400

hare, snowshoe, Lepus americanus, 54

harrier, hen, Circus cyaneus, 323, 324, 325, 327, 329, 330, 368, 401

Harris, 77

hawthorn, Crataegus monogyna, 115

hazel, Corylus avellana, 267, 270, 277

heath, cross-leaved, Erica tetralix, 310

heather, Calluna vulgaris, 22, 23, 39, 276-7, 290, 412-13

beetle, 308, 330, 396, 414

bell, Erica cinerea, 310

browning, 38, 62, 228, 308, 339-40

check, 154

habitat, 82, 119, 188-9, 285, 287, 318, 330, 358

in diet, 10, 38-40, 83-4, 113-5, 156, 230-1, 291, 289, 294-5, 297-8, 312

moorland, 9-11, 21, 39-40, 48, 50, 51, 64, 66, 109-15, 229, 280, 281-2, 284, 287-8, 359

management, 34, 38, 47, 118-19, 126, 282, 377, 387-406, 388, 390-2, 392-3, 394-7, 399

nutritive value, 39, 184, 228, 230, 261, 289, 300-3, 301-2, 305, 310

heath hen, Tympanuchus cupido cupido, 11

Hebrides, Inner, 78, 110, 245

Outer, 31, 54, 77-8, 110, 222, 245, 270

hedgehog, Erinaceus europaeus, 66

Herefordshire, 109

Highlands, Scottish, 78, 260, 262, 265, 267-8, 332, 336, 396, 403

holly, Ilex aquilifolium, 10, 270

hormones:

effect on behaviour, 230, 233

effect on plumage, 247-50, 248-9, 257-8

horsetail, Equisetum, 83-4, 156

Hungary, 269

hybrid, 31, 58, 164, 165

Iberia, 146

Ice Age, 8-9, 167, 263, 269-70, 276, 279, 284, 288

Iceland, 33, 49-50, 69, 76, 78, 81, 83, 87-9, 91, 182, 187, 222, 250, 304, 319-20, 365, 373

Hrisey, 49-50, 82, 84, 88, 93-4, 98-9, 196, 228, 242-3

Husavik, 93

inbreeding, 44, 108, 152

Indian, American, 381

Inshriach, 150

insulation, 252

introductions, 33, 77, 99, 320, 331

Inuit, 253, 380

Invercauld, 52, 160-1

Inverness, 402

invertebrates, 88, 312-16

Italy, 97, 269

Japan, 37, 69, 76, 82, 92, 384

juniper, Juniperus communis, 113, 149, 152, 182, 265-6, 278, 398, 407, 410

Jura, 77, 278

Kazakhstan, 23, 31-2, 38, 49-50, 61, 66, 77, 111, 239

Kent, 109

Kerloch, 27, 39, 43-6, 50-3, 56-60, 189, 195, 216-19, 222, 226-33, 364, 394-7

Kielder, 54

Kintyre, 277

Kinveachy, 332

kite, black, Milvus migrans, 322, 324

Kola, 76, 208

Kuriles, 80

Labrador, 76-7, 87, 413

Lake District, 77, 265

Lammermuirs, 279

Langholm, 329-30, 368, 378, 401

larch, Larix, 10, 114, 154-7, 183, 309, 310, 377

Laurentia, 259

Leadhills, 397

Leicestershire, 109

lek, 20, 174, 237, 287, 385, 412

lime, Tilia, 266-7

Loch Laggan, 406

Loch Lomond, 167

Lochnagar, 33, 81, 221

Mala, 381

marten, pine, Martes martes, 318, 319, 320, 321, 324, 331, 333

and capercaillie, 331-4

meadowsweet, Filipendula ulmaria, 38

Meall Odhar, 91, 94, 98-9

metapopulation, 167

Midlands, 277

migration, 42-6, 87-8, 164, 174, 225

Millden moor, 383

Minnesota, 33, 202

Misty Law, 64, 401

Molophilus ater (crane-fly), 313

Monaco, 269

Mongolia, 32, 77

Montgomery, 109

Moorhouse, 283, 363

Moran effect, 371

Moray, 77, 167, 384

moult, 158, 211, 235-49, 258

mouse-ear, Cerastium, 84, 309

movements, 42-6, 87-8, 122-4, 1-62-5, 367, 386

muirburn, 282, 285

effects on grouse densities, 46, 49, 64, 375

effects on habitats, 282-3, 376, 389, 392, 394

Mull, 46, 77, 218, 221, 245, 396, 403

myrtle, bog, Myrica gale, 129, 274

names, origins of, 13-20, 18-19

Nearctic (New World), 1-4, 2-3, 8, 12, 162

Netherlands, 412

North Atlantic Oscillation, 124, 147, 347-8, 349, 353

North Pole, 77

North York Moors, 48

Norway, 31, 33, 38, 52, 54-5, 57-60, 69, 76, 82, 86, 89, 147, 163, 174, 182, 195, 224, 234, 238-9, 251, 257, 263, 278, 288, 313, 324, 334, 343-4, 365, 376, 378, 384, 387

Dovrefjell, 49, 213, 221-2, 227-8, 231, 233

Hitra, 238-9

Karlsøy, 62, 336

Tranøy, 49-50, 52-3, 57-8, 62, 64, 313

Varaldskogen, 152, 164

Nottinghamshire, 109 nutrition, 289-316

effect on breeding success, 308, 311, 312-16

oak, Quercus, 157, 266-7, 277

oat, Avena, 157, 270

Orkney, 77-8, 277, 324

owl, short-eared, Asio flammeus, 66

Palearctic (Old World), 1-4, 2-3, 8, 162, 334

parasites, 330, 341, 367

effects on breeding success, 62, 64, 336, 340, 374-5

roundworm, Ascaridia compar, 334

tapeworms, Hymenolepis microps & Paroniella urogalli, 334, 336-7

threadworm, caecal, Trichostrongylus tenuis, 42, 66, 195, 257, 336-9, 338, 340, 364, 373-5

tick, sheep, Ixodes ricinus, 66, 100, 342-6, 343, 400, 414

peat, 48, 60, 268-75, 273, 278-81, 284, 288

pectinations, 254

Pennines, 48, 110, 113, 124, 131, 281

Perthshire, 118, 143-4, 147, 156, 167

pheasant, Phasianus colchicus, 8, 126, 188, 344

Philaenus spumarius (spittle bug), 313

philopatry, 45, 108

Pinega, 138

pine, Pinus, 10-11, 266-70, 276-7, 308, 377, 398, 407, 409

Arolla, P. cembra, 155

lodgepole, P. contorta latifolia, 118, 154, 377

Scots, P. sylvestris, 187, 267

habitat, 119, 148, 278, 287-8, 399, 407, 414

in diet, 113, 155-6, 291, 303

plumage, 184-7, 252, 258

pollution, 65, 100, 412-14

Polytrichum (moss), 156

population, 349, 355, 358, 369, 376, 386-7, 414

decline, 11, 361-3, 374-5, 379, 384-5, 398

density, 308, 382-7, 395-7

fluctuations, 12, 347-79, 351-2, 362

breeding success during, 365, 366

causes of, 371-5

effects of habitat on, 375-8

effects of spatial organisation, 366-8

effects of weather on, 350, 353, 354, 355, 378-9

Portugal, 32

prairie-chicken, Attwater’s, Tympanuchus cupido attwateri, 11

greater, Tympanuchus cupido, 3, 8

lesser, Tympanuchus pallidicinctus, 3, 8

prairie grouse, 2, 3, 4, 8, 10

predators, 308, 312, 321-3, 324-35, 333, 346, 358, 367-9, 372-3, 378-9, 400, 412

control of, 55, 64-5, 319, 324, 329-32, 346, 375-6, 378, 400, 401-3, 402

effect on breeding success, 329, 333-4, 335, 368

pine marten, Martes martes, 331, 332-6, 333, 335

predator-prey relationships, 317-20, 372-3

raptors, 318-24, 322-3, 326-8, 332, 373, 403, 414

gyrfalcon, Falco rusticolus, 93-4, 97, 242, 318-19, 373

persecution, 329-30, 368, 378, 400, 401-2

Priestlaw, 64

ptarmigan, Lagopus, 2, 4, 8, 10, 13

rock, Lagopus mutus, 6, 15, 293, 326, 327, 411

behaviour, 68-9, 74, 91, 314

calls, 174, 175, 176-9, 218-22, 226

feeding, 180-3, 304, 308, 310

parental care, 191-2, 193-4, 196

predator avoidance, 190

preening, 184

resting, 187

roosting, 188-9

in snow, 197-210, 199, 205, 207-8

sexual, 179, 180

social, 178, 180, 189-90

territorial, 173-7, 178, 213-16, 221, 222-7, 223, 232-4, 244

diet, 83-6, 85, 98, 291, 300, 304, 307, 309

distribution, 33, 67, 76-80,

enemies, 93-4, 99-100, 190-1, 318-46, 368-9, 373-4, 412

growth, 314

habitat, 36, 37, 79-80, 81, 82-3, 90, 259-61, 269, 284-8, 285, 377-9

metabolism, 86-7

movements, 87-9

plumage, 68-9, 70-3, 235-44, 246-50, 248-9, 251, 253, 254-5, 258

population:

density, 36, 88-9, 91, 92, 212-5, 222-4, 242, 348-9, 355

fluctuations, 91, 350-4, 352-3, 365-9, 366, 372-3, 379

size, 78

reproduction:

breeding success, 98-9, 230-2, 311, 354, 365, 366, 368

chicks, 74, 75, 79, 86-8, 99, 181, 226, 284, 294

eggs, 26, 69, 74, 95-8, 97

incubation, 86, 191, 194-6, 226, 242, 250, 257

nests, 85, 96

sex ratio in spring, 93-5

shooting, 99, 233, 381, 384

size, 68-9, 314

survival, 91-3, 92, 311, 366, 387

territory, 93-4, 211-12, 216-17, 218, 220, 223-34

weight, 24-6, 68-9, 86

white-tailed, Lagopus leucurus

behaviour, 78, 86, 199, 203

evolution, 2, 8, 9-10

habitat, 78-9

plumage, 237-8, 246, 254

size, 69, 158

willow, Lagopus lagopus

behaviour, 31, 40, 41

calls, 174-9, 175, 218-22, 226

feeding, 180-3, 303-7, 312-13

parental care, 191-6

predator avoidance, 190

preening, 184

resting, 187

roosting, 188-9

in snow, 197-210, 198, 205, 207-8

sexual, 179-80

social, 178, 180, 189-90

territorial, 173-8, 211-16, 220-7, 232-4, 244

diet, 38-40, 303-7, 312-13

distribution, 31-3, 32, 269

enemies, 64-6, 190-1, 318-46, 374, 413

growth, 27, 29, 42

habitat, 34-8, 35, 36, 37, 67, 269-70, 287

metabolism, 40, 41, 251

movements, 42-4

plumage, 21, 22, 31, 235-9, 244, 246-58, 248-9

population:

decline, 54, 55

density, 36, 49-50, 212-13

fluctuations, 52-3, 365, 378

size, 33

reproduction:

breeding success, 63 64, 195, 230-2, 365

chicks, 27, 40, 42, 74, 226

eggs, 27, 58-60, 61

incubation, 41, 42, 59, 194-5, 226, 250

nesting, 58-9, 62

sex ratio in spring, 57

shooting, 55, 56, 381, 384

size, 68

survival, 42, 56, 57, 366, 387

territory, 57, 211-12, 216-17, 218-20, 223-4, 226-8, 230-2, 234

weight, 23, 24-6, 62, 68

Pyrenees, 82, 89, 146, 163-4, 385, 412

Rannoch Moor, 265, 268

rape, Brassica napus, 157

recreation, 410-12

reintroduction, 9, 133, 147, 172, 324, 332

return rates, 56, 91-3, 92

rhododendron, Rhododendron ponticum, 110

Rickarton Moor, 52, 59, 195, 218-19, 222, 224, 227, 308, 366, 370, 372, 387, 398

rodents, 52-4, 318, 320, 330, 332-5, 333, 368

roost, 188-9, 197-210, 303

Rothiemurchus, 288, 410

rowan, Sorbus aucuparia, 115, 398

Rum, 77, 277

Russia, 23, 27, 32-3, 43, 54, 58-60 67, 76, 78, 81, 87, 99, 147, 190, 201, 254, 286, 320, 337, 381, 384-5, 407

Ilexa basin, 138, 307, 319, 324, 332-3

Lake Ilmen, 324

Pechora-Illych reserve, 163, 332

Pinega, 138

Timansk, 39-40

Urals, 76, 113, 147-8, 163, 168

saxifrage, purple, Saxifiaga oppositifolia, 84, 223

Scandinavia, 57, 109, 146-7, 151, 260, 263, 286, 336, 413

Scarba, 77

sex ratio, 94-5, 314

Sgribhis Bheinn, 33

Shetland, 278

Shropshire, 109

Siberia, 1, 8, 23, 29-32, 40, 53, 67, 69, 76-7, 109, 147, 197, 203, 206, 208, 252

Sinkiang, 77

size, 68, 135-6, 157-8, 314

ski development, 100, 318, 320, 368, 369, 410-12

Slovenia, 110

snares, 65, 170-1, 384, 400

snow, 24, 43, 59, 220, 224-6, 286, 355

effect on plumage, 239-40, 246

feeding in, 10, 38, 182-4, 308

roosts, 1, 40, 69, 197-210, 198-200, 205, 207-8, 252, 315

shoes, 253

Snowdonia, 265

soil, 57, 94, 112, 270-81, 271, 284-8, 311, 377, 410

Somerset, 109

sorrel, sheep’s, Rumex acetosella, 38, 309

Southern Uplands, 77-8, 110, 260

Spain, 146, 148

sparrowhawk, Accipiter nisus, 66, 190, 319, 322, 325, 332

Special Protection Area (SPA), 145-6

Speyside, 154, 167, 332, 345, 396

spruce, Picea, 10, 113

Norway, P. abies, 148, 155, 187, 189, 303

Sitka, P. sitchensis, 113, 114-5, 118, 154-6, 159, 291, 377-8, 391, 409, 410

Spyhill, 59, 183, 188

Staffordshire, 109

status, 11-12

Stirling, 147

stoat (short-tailed weasel), Mustela erminea, 99, 131, 321, 325, 334

Strathavon, 164

Strathmore, 60

study areas, xi, xii

subspecies, 31, 76, 109, 146, 222

Surrey, 109

survival, 314-16

Sussex, 109

Sutherland, 77, 81, 147, 267, 281

Svalbard, 33, 41, 67-8, 76-7, 86, 88, 91-2, 95, 97-9, 196, 201, 228, 231-4, 242, 251, 260, 308

Sweden, 33, 49, 52-3, 63-4, 76, 78, 110, 113, 147, 165, 287, 312, 333-4, 373, 376, 384

systematics, 31, 76, 109, 146

Taymouth, 147

temperature, 162-3, 207, 353, 354, 355

territory, 137, 164, 241, 339-40, 358-66, 360, 370, 374-5, 390

Thames, 269

toxins, plant, 300-4

Tweed Valley, 148

Uist, North, 270

South, 277

Unimak, 32

Urals, 76, 113, 147-8, 163, 168

Virginia, 269

Wales, 23, 31, 33, 48, 78, 100, 108-10, 116, 131, 245, 260, 267, 281-2, 287-8, 324, 342-3, 377, 396

weasel, Mustela nivalis, 99, 131, 324, 325

whortleberry, bog, Vaccinium uliginosum, 312, 407

willow, Salix:

effect of grazing, 94, 99

habitat, 38, 82, 265, 267, 270, 398

in diet, 38-40, 57, 83-4, 88-9, 113-14, 303-7, 306

Wiltshire, 109

wires, 65, 152, 410-12, 411

woodland grouse, Bonasa, 2, 4, 8, 154, 162, 174

Worcestershire, 109

Yakutia, 25

Yew, Taxus baccata, 270

Yorkshire, 277, 281