INDEX

Aché Indians, hunting by men of, 93–98

Adoptive mothers, lactation by, 51

Adultery laws, confidence in paternity and, 37–38. See also Extramarital sex

African elephants, menopause and, 125

African hunting dogs: infanticide and, 73

meat provisioning and, 92

Aging, 109–10

accidents and bad conditions and, 113–14

animal signals indicating, 130

human signals indicating, 135, 142

kingpin principle of, 114–16

repair and maintenance and, 109–13

repair and reproduction and, 112–13

signals indicating, 130, 135, 142

Albatrosses: “divorce” rates and, 135

eggs of, 108

parental care and, 17

Alexander, Richard, 71–73, 75

Alligators, parental care and, 23

American men, role of, 101–2

Amphibians: external fertilization and, 21

parental care and, 16

vivipary and, 22

Anatomical adaptations, natural selection and, 10–13

Andaman Islands, fat in women of, 138

Andelman, Sandy, 74–75

Androgens, 45

male pseudohermaphrodites and, 47

Animal signals, 128–34

age indicated by, 130

auditory, 129

behavioral, 129

in birds, 129, 130–31

electrical, 129

experimental study of, 130–31

Fisher’s runaway selection model of, 131–32

gender indicated by, 130

male sexual ornamentation and, 140

natural selection and, 128–29

olfactory, 129

sexual maturity indicated by, 130

truth in advertising theory of, 133–34

Zahavi’s handicap principle of, 132–33

Antelope, mating systems of, 7

Anticontraceptive theory, of concealed ovulation, 70–71

Ants, signals and, 129

Apes: aging and, 113–14

humans compared to, 8–10

ovulatory signs and, 78

penis of, 144

promiscuous mating system of, 82, 83

sexual distinctiveness and, 47–48. See also Bonobos; Chimpanzees; Gibbons; Gorillas; Orangutans

Archaeopteryx, 57

Arctic Indian men, hunting by, 95

Arctic seals, lactation and, 59

Asian elephants, menopause and, 125

Auditory signals, 129

Axillary hair, sexual maturity in humans indicated by, 135, 142

Baboons: ovulatory signals and, 64, 78, 82, 130

sex confined to estrus and, 64

Baleen whales, eggs of, 108

Bandicoots, pregnancy and, 58

Barbary macaques: public sex and, 3

sex confined to estrus and, 64–65

Bats, mating systems of, 7. See also Dyak fruit bats; Hammer-headed fruit bats

Battle of the sexes. See under Parental care

Behavioral signals, 3, 129

Better-genes-through-cuckoldry theory, of female sexual receptivity, 70

Big-bang reproduction (semelparity), 7, 106–7

Bipotential gonad, 44, 45

Birds: coparenting and, 29–34, 56

evolutionary commitment of, 55–58

internal fertilization and, 22, 55–56

mating systems of, 7

pair bonds formed by, 7–8

parental care and, 7, 16, 17, 55–58, 69

on Rennell Island, 122–23

sex for fertilization and, 66–67

sex-role-reversal polyandry and, 16, 26–29, 36, 57

signals and, 129, 130–31

Body fat. See Fat

Body signals. See Signals

Bonobos (pygmy chimpanzees): humans compared to, 8, 9

menopause and, 125

promiscuous mating system of, 8, 9, 69, 78–79

sex for fun and, 3

Bowerbirds, golden crest of, 132, 137

Breast-feeding. See under Lactation

Breasts: hormones and development of, 47–50

as signal in human females, 135, 136, 138–39, 141

Bride prices: confidence in paternity and, 37

women’s desirability and, 141

Brown, James, 133–34

Brush turkeys, parental care and, 17

Buttocks, as signal in human females, 138, 139

Century plants, big-bang reproduction and, 107

Chaperoning, confidence in paternity and, 38

Chickens, large eggs of, 27

Childbirth, risk of mother’s death and, 117–18

Child care. See under Parental care

Chimpanzees, 13

humans compared to, 8, 9

infanticide and, 73

lack of paternal care and, 2–3

menopause and, 125

ovulatory signals and, 78, 80, 81, 82

penis of, 144

private sex (consortship) and, 3

promiscuous mating system of, 9, 69, 78–79, 82, 83

public sex and, 3

pygmy, see Bonobos

Chromosomes, 43–44

Circumcision, female. See Clitoridectomy

Clitoridectomy, confidence in paternity and, 38

Clitoris, 45

Comparative method, concealed ovulation studied by, 77–88

Concealed ovulation, 5, 64, 67, 68–88

anticontraceptive theory of, 70–71

comparative method studying, 77–88

daddy-at-home theory of, 71–73, 75–76, 79, 84, 86–87

detecting ovulation and, 63–64

evolution of, 80–88

female receptivity and, 64–65

harems and, 84, 85

hormonal test kit and, 63–64

many-fathers theory of, 71, 73–75, 76, 79, 84

mating systems and, 79–80

paternity confused by, 71, 73–75

prostitution theory of, 70

theories explaining, 70–88

Confessions of Felix Krull, Confidence Man (Mann), 88

Confidence of parenthood: concealed ovulation and, 74–75

daddy-at-home theory of concealed ovulation and, 71–73, 75–76

genetic testing and, 60–61, 76

male lactation and, 43, 60–61

many-fathers theory of concealed ovulation and, 71, 73–75, 76

parental care and, 20, 24–25, 30, 37–38, 56

Conflict of interests: between closely related people, 100–101

in hunter-gatherer societies, 100

Consortship, chimpanzees and, 3

Coparenting: birds and, 29–34, 56

daddy-at-home theory and, 72

humans and, 29–30, 38

male lactation and, 53–54

sexual receptivity and, 68–70

Couples, humans forming, 4, 5–6

Cows, hormones and lactation and, 48

virgin cows and, 49

Daddy-at-home theory, of concealed ovulation, 71–73, 75–76, 79, 84, 86–87

Darwin, Charles, 10

Deer, antlers of, 133–34

Dihydrotestosterone, 45

male pseudohermaphrodites and, 47

Dinosaurs, 57

Divorce, 102

human signals and, 135

Dogs: mammary glands and, 48

sex for fertilization and, 66

signals and, 129. See also African hunting dogs

Dolphins, sex for fun and, 3

Dyak fruit bats, spontaneous male lactation and, 52, 54, 60

Echidnas, internal fertilization and, 22

Eggs: larger than sperm, 21

menopause and depletion of, 107–9, 115

Electrical signals, 129

Electric fishes, signals and, 129

Elephants: eggs of, 108

menopause and, 125

pregnancy and, 58

Enzymes, male/female organ development and, 46

Epididymis, 45

Estrogen, lactation and, 49–50

Estrus, mammals confining sex to, 64–65. See also Fertilization, mammals having sex for

Evolution: of concealed ovulation, 80–88

human sexuality and, 10–13. See also Natural selection

Evolutionary commitment, 54–55

birds’ reproductive strategy and, 55–58

mammalian reproductive strategy and, 58–60, 62

External fertilization, 21

alternative opportunities foreclosed by child care and, 23

in fish, 13

humans and, 61

obligate parental investment and, 21

parental care and, 25–26

Extramarital sex, 5

biological significance of, 98–99

confidence of parenthood and, 76

hunting and, 98–100

recreational sex and, 69–70

as risky, 68

sex appeal and, 141

sex differences in attitudes toward, 38–39

Extra-pair copulations, of Pied Flycatchers, 33–34

Facial beauty, as human signal, 137–38

Fallopian tubes, 45

Fat, as signal in human females, 135, 138–39, 141

Female menopause. See Menopause

Females (human): breasts and, 135, 136, 138–39, 141

buttocks and, 138, 139

extramarital sex and, 38–39

facial beauty and, 137–38, 141

fat and, 135, 138–39, 141

fertility decline in, 103, see also Menopause

hormones and breast development and, 49–50

lactation by, 23, 24, 35–36, 49–50

parental care and, 4, 21–24, 25, 29–30, 35–38, 102

plant foods gathered by, 91, 93–94, 95, 100

sexual maturity of, 142

sexual ornamentation and, 138–41. See also Concealed ovulation; Lactation; Menopause; Sexual receptivity (female)

Females (mammalian): duration of fertility of, 105–6, 115

lactation and, 23, 24, 49, 50

menopause and, 105–7, 124–25

parental care and, 7, 15–16, 21–34, 55–57

sex-role-reversal polyandry and, 26–29

signals and, 130, 131–34

Fertilization, mammals having sex for, 3, 64–65, 66–67. See also External fertilization; Internal fertilization

Fish: external fertilization and, 13, 21

male role and, 7

mating systems of, 7

parental care and, 7, 16, 25–26

vivipary and, 22

Fisher, Ronald, Sir, 131–32, 144

Fisher’s runaway selection model of signals, 131–32, 144

Flycatchers. See Pied Flycatchers

Ford, Henry, 114–15

Foreclosed opportunity, parental care and, 20, 23–24, 35–37, 38, 56

Frogs: mating systems of, 7

parental care and, 7, 25–26

Fun, sex for. See Recreational sex

Genes, 43, 47

males passing to offspring, 24. See also Chromosomes; Natural selection

Genetic testing, confidence of parenthood and, 60–61

Gibbons: couples and, 4

as monogamous, 78, 82

parental care and, 3, 54, 69

paternal care and, 3

sex confined to estrus and, 65

sexual distinctiveness and, 48

Glans penis, 45

Glass frogs, parental care and, 26

Goats (female), lactation in virgin: hormones and, 49

“milking” and, 50

Goats (male), lactation and: hormones and, 49

spontaneous lactation and, 42, 51–52

Gorillas: harems and, 5, 78, 82

head shape indicating sex and, 130

humans compared to, 9

infanticide and, 73, 74

ovulatory signals and, 78, 80, 81

paternal care and, 3

penis of, 143, 144

sex and parental care and, 69

sexual distinctiveness and, 48

sexuality of, 9

sexual maturity indicated and, 130

Grandmothers, extended role of, 119–20

Great apes. See Apes

Great Tits, breast stripe as signal and, 130–31

Growth hormone. See Somatotropin

Guinea pigs, lactation and, 59

hormones and male lactation and, 49

Hadza, foraging by women of different ages and, 119–20

Hammer-headed fruit bats, lek mating system of, 7, 8

Hamsters, duration of fertility of, 115

Handicap principle. See Zahavi’s handicap principle of signals

Harems: concealed ovulation and, 84, 85

humans and, 4, 5–6, 37

mammals and, 5

primates and, 78, 79, 80, 82, 83

Hawkes, Kristen, 93–98, 119–20

Hazda Indians, hunting by men of, 93, 95–96

Head shape, gender indicated by, 130

Herring Gulls, signals and, 130

Hill, Kim, 93

Hoofed mammals, lack of paternal care and, 2

Hormones: lactation and, 48–50

male lactation and, 47–50

sex differences and, 47–49

Howler monkeys, concealed ovulation and, 82

Hrdy, Sarah, 71, 73–75

Human sexuality: apes compared to, 8–10

apparent inefficiency of, 67–68

continuous practice of sex and, see Sexual receptivity (female)

coparenting and, 29–30, 38

evolutionary biology and, 10–13

extramarital sex and, 5, 38–39, 68, 69–70

harems and, 4, 5–6, 37

internal fertilization and, 21–23

long-term sexual partnerships and, 4, 5–6

mammalian sexuality compared to, 1–10

monogamy and, 5–6, 82, 83

ovulatory signals and, see Concealed ovulation; polyandry and, 6

polygyny and, 6, 37

premarital sex and, 5

private sex and, 4

promiscuity and, 78, 79

proximity to sexual partnerships of others and, 4

sequential monogamy and, 6. See also Concealed ovulation; Females (human); Males (human); Menopause; Parental care (human); Recreational sex; Sexual receptivity (female)

Human signals, 128–30, 134–46

age, 135, 142

facial beauty, 137–38, 141

handicap principle of, 133, 144–45

male voice, 127–28

men’s muscles, 135, 136–37

natural selection and, 128–29

penis, 142–46

runaway selection model of, 131–32, 144

sex appeal and, 135, 141

sexual maturity, 128–29, 135, 142

sexual ornamentation, 138–41, 142–46

truth in advertising principle of, 133–35, 136–39

women’s breasts, 136, 138–39, 141

women’s fat, 135, 138–39, 141

Hunter-gatherers: conflict of interests and, 100

lactational amenorrhea and, 36

weaning and, 59

women and, 91, 93–94, 95, 100. See also Hunters, men as

Hunters, men as, 91–101

Aché Indians and, 93–98

extramarital sex and, 98–100

Hazda Indians and, 93, 95–96

meat provisioning and, 91–93

as providers, 98–99, 100

as show-offs, 98–100

women’s jobs versus, 91, 93–94, 95, 100

Hurtado, A. Magdalena, 93

Hutterite women, babies produced by, 103

Imprisonment of women, confidence in paternity and, 38

Infanticide: conflict of interests leading to, 100–101

many-fathers theory and, 73–75, 76

Infibulation, confidence in paternity and, 38

Insects, mating systems of, 7

Interior vagina, 45

Internal fertilization, 13

alternative opportunities foreclosed by child care and, 23–24

birds and, 22, 55–56

confidence of parenthood and, 25

evolutionary commitment of mammals to, 58–60

obligate parental investment and, 21–23

Inuit (Eskimo) men, hunting by, 95

Investment in young: external fertilization and, 61

male lactation and, 43

parental care and, 20–23, 35, 38, 56

Ismail the Bloodthirsty, Emperor of Morocco, 37

Jacanas (lily-trotters), sex-role-reversal polyandry and, 26–29, 57

Jackrabbits, lactation and, 59

Jones, Nicholas Blurton, 93

Jost, Alfred, 44

Kangaroos, parental care and, 17

Kaplan, H., 93

Ketengban tribe, phallocarps in, 142–43

Killer whales, menopause and, 125

Kingpin principle, aging and, 114–16

Kodric-Brown, Astrid, 133–34

Labia majora, 45

Labia minora, 45

Lactational amenorrhea, 36

Lactation (female), 23, 35–36

evolutionary commitment to, 58–60, 62

hormones and, 48–50

humans and, 23, 24, 35–36, 49–50

infanticide terminating, 73

mammals and, 23, 24, 49, 50

mammary gland development and, 48

nipple stimulation and, 50–51

origin of, 58

pregnancy and, 58

species differences and, 59

weaning and, 59. See also Lactation (male)

Lactation (male), 41–62

advantages of, 61–62

conditions favoring, 60–61

evolutionary commitment and, 58–60

hormones and, 48–50

male roles and, 53–54

nipple stimulation and, 50

physiological feasibility of, 43–52

pregnancy necessity and, 42, 49

psychological obstacles to, 62

spontaneous lactation and, 42, 51–52, 60

starvation recovery and, 51

technological possibility of, 61–62

Langur monkeys, concealed ovulation and, 79

Lek mating system, of hammer-headed fruit bats, 7, 8

Life span, female reproduction and, 105

Lily-trotters. See Jacanas

Lions: infanticide and, 73

parental care and, 2–3, 53–54

Long-Tailed Widowbirds, tails as signal and, 130, 132

Macaques, ovulatory signals and, 82. See also Barbary macaques; Rhesus macaques; Stumptailed macaques

Male lactation. See Lactation (male)

Male pseudohermaphrodites, 46–47

Males (human): American men, 101–2

divorce and, 102

duration of fertility of, 103, 115

facial beauty and, 137–38

mammary gland development and, 48–50, see also Lactation (male)

menopause and, 5

muscles and, 135, 136–37

in New Guinea, 90–91, 95

parental care and, 4, 6, 15, 16, 19, 21, 22, 24, 25, 29–30, 35–38, 71–73, 75, 102

penis of, 142–46

as providers, 98–99, 100, 102

role of, 89–102

sexual maturity of, 128–29, 135, 142

sexual ornamentation and, 140, 142–46

as show-offs, 98–100, 102

voice change and, 127–28. See also Extramarital sex; Hunters, men as; Lactation (male)

Males (mammalian): duration of fertility of, 103, 105–6, 115

infanticide and, 73–74

mammary gland development and, 48, see also Lactation (male)

meat provisioning and, 92

menopause and, 106–7

parental care and, 2–3, 7, 15, 21, 24, 25–34, 55–58

post copulatory desertion and, 23–25

role of, 2, 7, 15

sex-role-reversal polyandry and, 26–29

signals and, 130, 131–34, 140. See also Lactation (male)

Mammalian sexuality, 2–4

fertility time span and, 4

for fertilization, 3, 64–65, 66–67

harems and, 5, 78, 79, 80, 82, 83

human sexuality compared to, 1–10

male’s role and, 2, 7

mating systems, 7–8

menopause and, 4

one-night stands and, 5

ovulatory signals and, 3, 64

public sex and, 3. See also Females (mammalian); Internal fertilization; Males (mammalian); Parental care (mammalian)

Mammary gland development, hormones and, 47–50. See also Breasts; under Lactation

Mann, Thomas, 88

Mantises, sexual cannibalism and, 7, 11–12

Many fathers theory, of concealed ovulation, 71, 73–75, 76, 79, 84

Marmosets: as monogamous, 78

ovulatory signals and, 78

parental care and, 54

Marriage, by humans, 4

Marsupial mice, 8

menopause in males of, 106–7

sex for fertilization and, 66

Marsupials: internal fertilization and, 13

lactation and, 50, 58. See also Kangaroos

Mating systems: concealed ovulation and, 79–80

evolutionary changes and, 80–88

of primates, 78–79, 80–88

Men. See under Male

Menopause, 5, 10, 103–25

aging and, 109–13

burden of parental dependence of child and, 116–17

costs and benefits of, 116–20

definition of, 4

depletion and aging of egg supply and, 107–9, 115

extended role of postmenopausal women and, 118–20

importance to society of memories of old women and, 120–24

increase in expected life span and, 104–5

kingpin principle and, 114–16

in mammals, 106–7

more children produced by, 116–25

risk of death in childbirth and, 117–18

risks in infants of older mothers and, 118

Menstrual cycle, lactational amenorrhea and, 36. See also Menopause

Mice: aging and, 113

duration of fertility of, 106, 115

mammary glands in male, 48

repair versus reproduction and, 112–13. See also Marsupial mice

Midwife toads, parental care and, 16, 26

Missing Link, in concealed ovulation evolution, 80–88

Mixed reproductive strategy, of Pied Flycatchers, 33, 34

Møller, Anders, 77–88

Monkeys: concealed ovulation and, 82

monogamy and, 82

promiscuous mating system of, 82, 83. See also Night monkeys; Spider monkeys

Monogamy: concealed ovulation and, 79–81, 84–86

daddy-at-home theory and, 71–73, 75

of humans, 5–6, 82, 83

of primates, 78, 79, 82, 83

Monotremes: internal fertilization and, 22

lactation and, 58

Mormons, polygyny and, 37

Mullerian ducts, 45

Mullerian inhibiting hormone, 45

male pseudohermaphrodites and, 47

Muscles, as signal in human males, 135, 136–37

Napoleon the Third, 67

Natural selection, 104

behavior and, 16

concealed ovulation and, 87

lactation and, 60

male lactation and, 61

menopause and, 104

parental care and, 16–18

repair and reproduction and, 109–13

sexuality and, 10–13

signals and, 128–29

vertebrate limbs and, 87

New Guinea: bride prices in, 141

men’s role in, 90–91, 95

phallocarps in, 142–43

Night monkeys, concealed ovulation and, 79

Nizam of Hyderabad, 37

Noonan, Katharine, 71–73, 75

Nursing. See under Lactation

O’Connell, James, 93

Octopus, big-bang reproduction (semelparity) and, 7

Offspring, conflict of interest between parents and, 100–101

Olfactory signals, 3, 129

One-night stands, 5

On the Origin of Species (Darwin), 10

Opportunity. See Foreclosed opportunity

Orangutans: concealed ovulation and, 82

head shape indicating sex and, 130

humans compared to, 9

one-night stands and, 5

ovulatory signs and, 78

penis of, 143, 144

sexuality of, 9

Oring, Lewis, 27

Ostriches, parental care and, 17

Ovaries, 44

Ovulatory signals, 3, 130

evolutionary changes in, 80–88

in mammalian sexuality, 64, 77–78

in primates, 64, 77–78, 82, 130. See also Concealed ovulation

Parental care, 15–39

alternative opportunities foreclosed by child care and, 20, 23–24, 35–37, 38

battle of the sexes and, 18–20

confidence of parenthood and, 20, 24–25, 37–38, 38

male lactation and, 53–54

sex differences in obligate parental investment and, 20–23, 35, 38

Parental care (human), 2, 4, 6, 15, 34–38

alternative opportunities foreclosed by child care and, 23–24, 35–37

battle of the sexes and, 18–20

burden of long period of, 116–17

confidence of parenthood and, 24–25, 37–38

conflict of interests and, 100–101

coparenting and, 29–30, 38

females and, 4, 21–24, 25, 29–30, 35–38, 102

males and, 4, 6, 15, 16, 19, 21, 22, 24, 25, 29–30, 35–38, 71–73, 75, 102

polygyny and, 37

recreational sex and, 68–70

sex differences in obligate parental investment and, 21–23, 35

unwed mothers and, 6

Parental care (mammalian), 2–3, 7, 13, 15–34

alternative opportunities foreclosed by child care and, 23–24, 56

battle of the sexes and, 18–20

choices in, 16–18

confidence of parenthood and, 24–25, 56

coparenting, 29–34, 56

females and, 7, 15–16, 21–34, 55–57

lactation and, 53–54

male post-copulatory desertion and, 23–25

males and, 2–3, 7, 15, 21, 24, 25–34, 55–58

natural selection and, 16–18

sex and, 69

sex differences in obligate parental investment and, 22, 56

sex-role-reversal polyandry and, 26–29

Parenthood, confidence of. See Confidence of parenthood

Parricide, conflict of interests leading to, 100–101

Paternal care, 2–3. See also under Male

Peacocks, 8

tail of, 132

Penis, 44

internal fertilization and, 21

as signal in human males, 142–46

Penis shaft, 45

Phalaropes. See Wilson’s Phalaropes

Phallocarps, 143–44

Pheromones, 77

Pied Flycatchers: coparenthood and, 30–34

sex for fertilization and, 66–67

Pigeons: lactation and, 42, 56

small eggs of, 27

Pilot whales, menopause and, 107, 124–25

Placentals, lactation and, 48, 58

Platypus, internal fertilization and, 22

Plummage, as signal, 130

Polyandry, 36

humans and, 6

sex-role-reversal, 26–29, 36, 57

Tre-ba of Tibet and, 36, 37

Polygamy, harems and, 6

Polygyny: coparenthood in Pied Flycatchers and, 30–34

humans and, 6, 37

Porcupines, aging and, 113

Post-copulatory desertion by males, 23–25

exceptions to, 25–34

Pregnancy: duration of, 58

lactation and, 42, 49, 58

Premarital sex, humans and, 5

Primary female, polygyny in Pied Flycatchers and, 30–31, 32

Primates: concealed ovulation studied with, 77–88

family tree of, 80–88

mammary glands and, 48

mating systems of, 78–79, 80–88

ovulatory signals and, 64, 77–78, 82, 130

promiscuity of, 78–79, 80, 82, 83. See also specific primates, i.e. Apes; Baboons; Chimpanzees, etc.

Primordial gonad, 44, 45

Private sex, humans and, 4

Progesterone, lactation and, 49–50

Prolactin, lactation and, 48, 49

Promiscuity: concealed ovulation and, 84, 85

of humans, 79

of primates, 78–79, 80, 82, 83

Prostate gland, 44

Prostitution theory, of concealed ovulation, 70

Providers, men as, 98–99, 100, 102

Proximate mechanism, ultimate explanations versus, 108–9

Pseudohermaphrodites, 46–47

Pubic hair, sexual maturity in humans indicated by, 135, 142

Public sex, mammals and, 3

Pygmy chimpanzees. See Bonobos

Pythons, parental care and, 23

Rabbits, duration of fertility of, 115

Rats (male), mammary glands and, 48

Recreational sex, 4, 5, 10, 66, 67

coparenting and, 68–70

extramarital sex and, 69–70

sex-to-promote-family-values theory and, 68–69. See also Concealed ovulation; Sexual receptivity (female)

Rennell Island, consumption of fruits and seeds by birds and, 122–23

Repair and maintenance, 109–13

death and accidents and, 113

reproduction versus, 112–13

Repair and reproduction, natural selection and, 109–13

Reproductive strategy, 17

Reptiles: therapsid, 58

vivipary and, 22

Rhesus macaques, duration of fertility of, 106, 115

Rockefeller, Nelson, 67

Runaway selection model. See Fisher’s runaway selection model of signals

Saddleback tamarin monkeys, paternal care and, 3

Salmon, big-bang reproduction (semelparity) and, 7, 107

San (Bushmen): fat in women of, 138

men as hunters and, 95

Sandpipers. See Spotted Sandpipers

Scrotum, 45

Sea birds, pair bonds and, 8

Seahorses: male pregnancy and, 42

parental care and, 16

Seals. See Arctic seals

Secondary female, polygyny in Pied Flycatchers and, 31–32

Semelparity. See Big-bang reproduction

Seminal vesicles, 45

Senescence. See Aging

Sequential monogamy, humans and, 6

Sex appeal, 135

extramarital sex and, 141

as signal, 135, 141

Sex chromosomes, 43–44, 45, 47

Sex differences, hormones and, 47–49

Sex for fertilization. See Fertilization, mammals having sex for

Sex for fun. See Recreational sex

Sex in private, humans and, 4

Sex in public, mammals and, 3

Sex-role-reversal polyandry, 26–29, 36, 57

Sex-to-promote-family-values theory, of recreational sex, 68–69

Sexual cannibalism, 7, 11–12

Sexual evolution, 11–13

Sexual maturity, signals indicating, 128–29, 130, 135, 142

Sexual ornamentation: function of, 145

in men, 140, 142–46

in women, 138–41

Sexual receptivity (female), 5, 64–65, 67

better-genes-through-cuckoldry theory of, 70

coparenting and, 68–70

daddy-at-home theory and, 71–73, 75

extramarital sex and, 69–70

infanticide and, 73–75, 76

many-fathers theory and, 71, 73–75

Sexual signals. See Signals

Shore birds, sex-role-reversal polyandry and, 26–29, 57

Show-offs, men as, 98–100, 102

Siblicide, conflict of interests leading to, 100–101

Signals, 128–46. See also Animal signals; Human signals; Ovulatory signals

Sillén-Tullberg, Birgitta, 77–78, 88

Social systems, evolution and, 11

Somatotropin (growth hormone), 48

Songbirds, sex and parental care and, 69

Sperm: costliness of production of, 67

egg larger than, 21

Spider monkeys, ovulatory signs and, 78

Spiders: male role and, 7

sexual cannibalism and, 7, 11–12

Spontaneous male lactation, 42, 51–52, 60

Spotted Sandpipers: parental care and, 16

sex-role-reversal polyandry and, 26–29, 57

Starvation, lactation in men and recovery from, 51

Steatopygia, 138, 139

Steers, hormones and milk production and, 49

Steroids, 45

Sticklebacks, parental care and, 16, 26

Stumptailed macaques, spontaneous male lactation and, 52

Testes, 45

male pseudohermaphrodites and, 47

testosterone and, 45

Y chromosome and, 44, 45

Testosterone, 45

effects of, 47–48

male pseudohermaphrodites and, 46–47

Therapsid reptiles, lactation and, 58

Tigers, one-night stands and, 5

Toads, parental care and, 16. See also Midwife toads

Tortoises, eggs of, 108

Tre-ba, polyandry and, 36, 37

Truth in advertising theory of signals, 133–35, 136–39

Turtles, aging and, 113

Ultimate explanation, proximate mechanisms versus, 108–9

Unwed mothers, parental care from, 6

Uterus, 45

Vagina, 45

male pseudohermaphrodites and, 47

Vas deferens, 45

Vervets: concealed ovulation and, 74–75, 79

ovulatory signals and, 78

promiscuous mating system of, 78–79

Virgins as brides, confidence in paternity and, 37

Visual signals, 64, 78, 82, 130

Vivipary, 22

Voice change in males, as signal, 127–28

Weaning, in hunter-gatherer societies, 59

Whales. See Baleen whales; Killer whales; Pilot whales

Widowbirds. See Long-Tailed Widowbirds

Wilson’s Phalaropes, sex-role-reversal polyandry and, 16, 26–29, 36, 57

Witch’s milk, 49

Wolffian ducts, 45

Wolves: lack of paternal care and, 2–3

meat provisioning and, 92

parental care and, 54

Women. See Females (human)

X chromosome, 43–44

Y chromosome, 43, 44, 45

male pseudohermaphrodites and, 47

Zahavi, Amotz, 132–33, 144–45

Zahavi’s handicap principle of signals, 132–33, 144–45

Zebras: harems and, 5

paternal care and, 3