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Index
MAMMALS OF WASHINGTON
CONTENTS
ILLUSTRATIONS
INTRODUCTION
PHYSIOGRAPHIC PROVINCES OF THE STATE
DISTRIBUTIONAL AREAS
CLIMATE AND VEGETATION
LIFE-ZONES AND ECOLOGY
The Humid Subdivision of the Transition Life-zone
The Arid Timbered Subdivision of the Transition Life-zone
The Arid Grasslands Subdivision of the Transition Life-zone
The Upper Sonoran Life-zone
The Canadian Life-zone
The Hudsonian Life-zone
Table 1. Distribution of mammals in Washington by Life-Zones. A. Abundant. C. Common. R. Rare.
GEOLOGIC HISTORY OF WASHINGTON
THE FAUNAS
SPECULATION AS TO EMIGRATIONAL HISTORY OF THE MAMMALS
SPECULATION AS TO THE LATER DISTRIBUTIONAL HISTORY OF THE MAMMALS
EXPLANATION OF TREATMENT
CHECK LIST OF MAMMALS
CLASS MAMMALIA—mammals
ACCOUNTS OF SPECIES AND SUBSPECIES
Didelphis virginiana virginiana Kerr Opossum
Neurotrichus gibbsii
Gibbs shrew-mole
Neurotrichus gibbsii gibbsii (Baird)
Neurotrichus gibbsii minor Dalquest and Burgner
Genus Scapanus Pomel Moles
Scapanus townsendii (Bachman) Townsend mole
Scapanus orarius Coast mole
Scapanus orarius orarius True
Scapanus orarius yakimensis Dalquest and Scheffer
Scapanus orarius schefferi Jackson
Genus Sorex Linnaeus Long-tailed shrews
Sorex cinereus Kerr Cinereous shrew
Sorex cinereus cinereus Kerr
Sorex cinereus streatori Merriam
Sorex merriami merriami Dobson Merriam shrew
Sorex trowbridgii Baird Trowbridge shrew
Sorex trowbridgii trowbridgii Baird
Sorex trowbridgii destructioni Scheffer and Dalquest
Sorex vagrans Baird Wandering shrew
Sorex vagrans vagrans Baird
Sorex vagrans monticola Merriam
Sorex obscurus Merriam Dusky shrew
Sorex obscurus obscurus Merriam
Sorex obscurus setosus Elliot
Sorex palustris navigator (Baird) Mountain water shrew
Sorex bendirii (Merriam) Bendire water shrew
Sorex bendirii bendirii (Merriam)
Sorex bendirii albiventer Merriam
Microsorex hoyi washingtoni Jackson Pigmy shrew
Genus Myotis Kaup Mouse-eared bats
Myotis lucifugus (Le Conte) Big myotis
Myotis lucifugus carissima Thomas
Myotis lucifugus alascensis Miller
Myotis yumanensis (H. Allen) Yuma myotis
Myotis yumanensis sociabilis H. W. Grinnell
Myotis yumanensis saturatus Miller
Myotis keenii keenii (Merriam) Keen myotis
Myotis evotis (H. Allen) Long-eared myotis
Myotis evotis evotis (H. Allen)
Myotis evotis pacificus Dalquest
Myotis thysanodes thysanodes Miller Fringe-tailed myotis
Myotis volans (H. Allen) Hairy-winged myotis
Myotis volans longicrus (True)
Myotis volans interior Miller
Myotis californicus (Audubon and Bachman) California myotis
Myotis californicus caurinus Miller
Myotis californicus californicus (Audubon and Bachman)
Myotis subulatus melanorhinus (Merriam) Small-footed myotis
Lasionycteris noctivagans (Le Conte) Silver-haired bat
Corynorhinus rafinesquii (Lesson) Long-eared bat
Corynorhinus rafinesquii townsendii (Cooper)
Corynorhinus rafinesquii intermedius H. W. Grinnell
Pipistrellus hesperus hesperus (H. Allen) Western pipistrelle
Eptesicus fuscus bernardinus Rhoads Big brown-bat
Lasiurus cinereus cinereus (Beauvois) Hoary bat
Antrozous pallidus cantwelli Bailey Pallid bat
Ursus americanus Pallas Black bear
Ursus americanus altifrontalis Elliot
Ursus americanus cinnamomum Audubon and Bachman
Ursus chelan Merriam Grizzly bear
Procyon lotor (Linnaeus) Raccoon
Procyon lotor psora Gray
Procyon lotor excelsus Nelson and Goldman
Martes caurina Merriam Western marten
Martes caurina caurina (Merriam)
Martes caurina origenes (Rhoads)
Martes pennanti (Erxleben) Fisher
Mustela erminea Linnaeus Ermine
Mustela erminea invicta Hall
Mustela erminea fallenda Hall
Mustela erminea olympica Hall
Mustela erminea streatori (Merriam)
Mustela erminea gulosa Hall
Mustela erminea murica (Bangs)
Mustela frenata Lichtenstein Long-tailed Weasel
Mustela frenata nevadensis Hall
Mustela frenata effera Hall-
Mustela frenata washingtoni (Merriam)
Mustela frenata altifrontalis Hall
Mustela vison energumenos (Bangs) Mink
Gulo luscus luteus Elliot Wolverine
Lutra canadensis (Schreber) River otter
Lutra canadensis pacifica Rhoads
Lutra canadensis vancouverensis Goldman
Enhydra lutris nereis (Merriam) Sea otter
Spilogale gracilis Merriam Civet cat or spotted skunk
Spilogale gracilis saxatilis Merriam
Spilogale gracilis latifrons Merriam
Mephitis mephitis (Schreber) Striped skunk
Mephitis mephitis hudsonica Richardson
Mephitis mephitis major (Howell)
Mephitis mephitis notata (Howell)
Mephitis mephitis spissigrada Bangs
Taxidea taxus taxus (Schreber) Badger
Vulpes fulva cascadensis Merriam Red fox
Canis latrans Say Coyote
Canis latrans lestes Merriam
Canis latrans incolatus Hall
Canis lupus fuscus Richardson Timber Wolf
Canis familiaris Linnaeus Dog
Felis concolor Linnaeus Cougar or mountain lion
Felis concolor oregonensis Rafinesque
Felis concolor missoulensis Goldman
Lynx canadensis canadensis Kerr Canadian lynx
Lynx rufus (Schreber) Bobcat
Lynx rufus fasciatus Rafinesque
Lynx rufus pallescens Merriam
Zalophus californianus (Lesson) California sea lion
Eumetopias jubata (Schreber) Steller sea lion
Callorhinus ursinus cynocephalus (Walbaum) Alaska fur seal
Phoca vitulina richardii (Gray) Hair seal or harbor seal
Genus Tamias Illiger Chipmunks
Tamias minimus Bachman Least chipmunk
Tamias minimus scrutator (Hall and Hatfield)
Tamias minimus grisescens (Howell)
Tamias amoenus Allen Yellow-pine chipmunk
Tamias amoenus caurinus (Merriam)
Tamias amoenus felix Rhoads
Tamias amoenus ludibundus (Hollister)
Tamias amoenus affinis Allen
Tamias amoenus canicaudus (Merriam)
Tamias amoenus luteiventris Allen
Tamias ruficaudus simulans (Howell) Red-tailed chipmunk
Tamias townsendii Bachman Townsend chipmunk
Tamias townsendii townsendii Bachman
Tamias townsendii cooperi Baird
Marmota monax petrensis Howell Woodchuck
Marmota flaviventris avara (Bangs) Yellow-bellied marmot
Marmota caligata cascadensis Howell Hoary marmot
Marmota olympus (Merriam) Olympic marmot
Citellus townsendii townsendii (Bachman) Townsend ground squirrel; sage rat
Citellus washingtoni Howell Washington ground squirrel; sage rat
Citellus columbianus (Ord) Columbian ground squirrel
Citellus columbianus columbianus (Ord)
Citellus columbianus ruficaudus Howell
Citellus beecheyi douglasii (Richardson) Beechey ground squirrel
Citellus lateralis (Say) Golden-mantled ground squirrel
Citellus lateralis tescorum (Hollister)
Citellus lateralis connectens (Howell)
Citellus saturatus (Rhoads) Golden-mantled ground squirrel
Sciurus griseus griseus Ord Western gray squirrel
Sciurus carolinensis hypophaeus Merriam Eastern gray squirrel
Sciurus niger Linnaeus subsp? Fox squirrel
Tamiasciurus hudsonicus (Erxleben) Red squirrel
Tamiasciurus hudsonicus streatori (Allen)
Tamiasciurus hudsonicus richardsoni (Bachman)
Tamiasciurus douglasii douglasii (Bachman) Douglas squirrel
Glaucomys sabrinus (Shaw) Northern flying squirrel
Glaucomys sabrinus oregonensis (Bachman)
Glaucomys sabrinus bangsi (Rhoads)
Glaucomys sabrinus columbiensis Howell
Glaucomys sabrinus latipes Howell
Glaucomys sabrinus fuliginosus (Rhoads)
Perognathus parvus (Peale) Great Basin pocket mouse
Perognathus parvus parvus (Peale)
Perognathus parvus lordi (Gray)
Perognathus parvus columbianus Merriam
Dipodomys ordii columbianus (Merriam) Ord kangaroo rat
Thomomys talpoides (Richardson) Northern pocket gopher
Thomomys talpoides devexus Hall and Dalquest
Thomomys talpoides columbianus Bailey
Thomomys talpoides aequalidens Dalquest
Thomomys talpoides wallowa Hall and Orr
Thomomys talpoides fuscus Merriam
Thomomys talpoides yakimensis Hall and Dalquest
Thomomys talpoides shawi Taylor
Thomomys talpoides immunis Hall and Dalquest
Thomomys talpoides limosus Merriam
Thomomys talpoides douglasii (Richardson)
Thomomys talpoides glacialis Dalquest and Scheffer
Thomomys talpoides tacomensis Taylor
Thomomys talpoides pugetensis Dalquest and Scheffer
Thomomys talpoides tumuli Dalquest and Scheffer
Thomomys talpoides yelmensis Merriam
Thomomys talpoides couchi Goldman
Thomomys talpoides melanops Merriam
Castor canadensis Kuhl Beaver
Castor canadensis leucodonta Gray
Castor canadensis idoneus Jewett and Hall
Onychomys leucogaster fuscogriseus Anthony Northern grasshopper mouse
Reithrodontomys megalotis megalotis (Baird) Western harvest mouse
Peromyscus maniculatus (Wagner) Deer mouse
Peromyscus maniculatus oreas Bangs
Peromyscus maniculatus hollisteri Osgood
Peromyscus maniculatus austerus (Baird)
Peromyscus maniculatus rubidus Osgood
Peromyscus maniculatus gambelii (Baird)
Peromyscus maniculatus artemisiae (Rhoads)
Neotoma cinerea Ord Bushy-tailed wood rat
Neotoma cinerea occidentalis Baird
Neotoma cinerea alticola Hooper
Synaptomys borealis wrangeli Merriam Northern lemming mouse
Phenacomys intermedius Merriam Heather vole
Phenacomys intermedius intermedius Merriam
Phenacomys intermedius oramontis Rhoads
Clethrionomys gapperi (Vigors) Gapper red-backed mouse
Clethrionomys gapperi saturatus (Rhoads)
Clethrionomys gapperi idahoensis (Merriam)
Clethrionomys gapperi cascadensis Booth
Clethrionomys gapperi nivarius (Bailey)
Clethrionomys californicus occidentalis (Merriam) California red-backed mouse
Microtus pennsylvanicus (Ord) Pennsylvanian meadow mouse
Microtus pennsylvanicus funebris Dale
Microtus pennsylvanicus kincaidi Dalquest
Microtus montanus (Peale) Montane meadow mouse
Microtus montanus nanus (Merriam)
Microtus montanus canescens Bailey
Microtus townsendii (Bachman) Townsend meadow mouse
Microtus townsendii townsendii (Bachman)
Microtus townsendii pugeti Dalquest
Microtus longicaudus (Merriam) Long-tailed meadow mouse
Microtus longicaudus halli Ellerman
Microtus longicaudus macrurus Merriam
Microtus richardsoni (De Kay) Water rat
Microtus richardsoni arvicoloides (Rhoads)
Microtus richardsoni macropus (Merriam)
Microtus oregoni oregoni (Bachman) Creeping mouse
Lagurus curtatus pauperrimus (Cooper) Sagebrush vole
Ondatra zibethicus (Linnaeus) Muskrat
Ondatra zibethicus osoyoosensis (Lord)
Ondatra zibethicus occipitalis (Elliot)
Rattus rattus (Linnaeus) Roof rat
Rattus rattus rattus (Linnaeus)
Rattus rattus alexandrinus (Geoffroy)
Rattus norvegicus norvegicus (Erxleben) Norway rat
Mus musculus Linnaeus, subsp? House mouse
Aplodontia rufa (Rafinesque) Mountain beaver
Aplodontia rufa rufa (Rafinesque)
Aplodontia rufa rainieri Merriam
Zapus princeps Allen Big jumping mouse
Zapus princeps trinotatus Rhoads
Zapus princeps kootenayensis Anderson
Zapus princeps idahoensis Davis
Zapus princeps oregonus Preble
Erethizon dorsatum (Linnaeus) Porcupine
Erethizon dorsatum epixanthum Brandt
Erethizon dorsatum nigrescens Allen
Myocastor coypus (Molina) subsp? Coypu, nutria
Ochotona princeps (Richardson) Pika or cony
Ochotona princeps cuppes Bangs
Ochotona princeps fenisex Osgood
Ochotona princeps brunnescens Howell
Lepus townsendii townsendii Bachman White-tailed jack rabbit
Lepus americanus Erxleben Snowshoe rabbit
Lepus americanus washingtonii Baird
Lepus americanus cascadensis Nelson
Lepus americanus pineus Dalquest
Lepus americanus columbiensis Rhoads
Lepus californicus deserticola Mearns Black-tailed jack rabbit
Sylvilagus nuttallii nuttallii (Bachman) Nuttall cottontail
Sylvilagus floridanus (Allen) subsp.? Florida cottontail
Sylvilagus idahoensis (Merriam) Pigmy rabbit
Cervus canadensis (Erxleben) Elk or wapiti
Cervus canadensis roosevelti Merriam
Cervus canadensis nelsoni Bailey
Odocoileus virginianus (Boddaert) White-tailed deer
Odocoileus virginianus ochrourus Bailey
Odocoileus virginianus leucurus (Douglas)
Odocoileus hemionus (Rafinesque) Mule deer and black-tailed deer
Odocoileus hemionus hemionus (Rafinesque)
Odocoileus hemionus columbianus (Richardson)
Alce americana shirasi Nelson Moose
Rangifer arcticus montanus Seton-Thompson Caribou
Bison bison oregonus Bailey Bison
Ovis canadensis Shaw Mountain sheep
Ovis canadensis canadensis Shaw
Ovis canadensis californiana Douglas
Oreamnos americanus (Blainville) Mountain goat
Oreamnos americanus americanus (Blainville)
Oreamnos americanus missoulae Allen
ORDER CETACEA Whales and porpoises
Berardius bairdii Stejneger Baird beaked whale
Mesoplodon stejnegeri True Stejneger beaked whale
Delphinus bairdii Dall Dolphin
Lissodelphis borealis (Peale) Right whale porpoise
Lagenorhynchus obliquidens Gill Striped porpoise
Grampus rectipinna (Cope) Killer whale
Grampidelphis griseus (Cuvier) Grampus
Globicephalus scammonii (Cope) Blackfish
Phocoena vomerina (Gill) Harbor porpoise
Phocoenoides dalli (True) Dall porpoise
Physeter catodon Linnaeus Sperm whale
Kogia breviceps (Blainville) Pigmy sperm whale
Rhachianectes glaucus (Cope) Gray whale
Balaenoptera physalus (Linnaeus) Finback whale
Balaenoptera borealis Lesson Sei or Pollack whale
Balaenoptera acutorostrata Lacépède Pike whale
Sibbaldus musculus (Linnaeus) Blue whale or sulphur-bottom whale
Megaptera novaeangliae (Borowski) Humpback whale
Eubalaena sieboldii (Gray) Pacific right whale
HYPOTHETICAL LIST
ADDENDA
BIBLIOGRAPHY
INDEX
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